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physher website  

Physher is a maximum likelihood-based method for estimating evolutionary rates and divergence times from genetic, amino acid, codon, and generic data. Phylogenetic trees should be open with Figtree from this website or Seqotron.

Environment Modules

Run module spider physher to find out what environment modules are available for this application.

System Variables

  • HPC_PHYSHER_DIR - installation directory
  • HPC_PHYSHER_BIN - executable directory
  • HPC_PHYSHER_LIB - library directory
  • HPC_PHYSHER_INC- include directory


If you publish research that uses physher you have to cite it as follows:

Fourment M and Holmes EC. Novel non-parametric models to estimate evolutionary rates and divergence times from heterochronous sequence data. BMC Evolutionary Biology 14:163, 2014.